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The story picks up where the first film left off, with Larry Daley (Ben Stiller) now running a successful museum in New York City. However, he's facing a new challenge: the American Museum of Natural History is expanding, and a new wing is being added, which will feature a collection of historical figures from around the world. The museum's curator, Jedediah (Owen Wilson), returns from the previous film, and Larry is tasked with overseeing the installation of the new exhibits.